For the past 40 years, anoline lizards have captured my curiousity, leading me into the investigation of their behavioral ecology. In particular, Anolis signalling systems have been a primary research interest. My belief that adaptive inference can best be drawn from free-ranging subjects has taken me to study sites throughout the Caribbean, Mexico, Central America, Hawaii, and southeastern US. Recent research questions have focused on the interactions of life history traits, morphology, social strategies, and display behavior of Anolis carolinensis as expressions of sexual selection.

EDUCATION

Doctor of Philosophy degree in Zoology,
  University of Oklahoma, 1969.
  Dissertation: Ethoecology and Display Analysis of Anolis nebulosus (Sauria, Iguanidae).
  Advisor, Charles C. Carpenter.

Master of Arts degree in Zoology,
  Southern Illinois University, 1964.
  Thesis: Food Habits of the Green Frog, Rana clamitans in Southern Illinois.
  Adviser, W. D. Klimstra.

Bachelor of Science degree in Biology,
  University of Redlands, 1962.  
  Research mentor, John Goodman. 

PROFESSIONAL EMPLOYMENT

2007-Present: Emeritus Status, Department of Biological Sciences, Virginia Polytechnic Institute and State University.

1977-2007: Associate Professor, Department of Biological Sciences, Virginia Polytechnic Institute and State University.

1999: Visiting Scientist, Smithsonian Institution (Natural History Museum), on sabbatical from VPI & SU.

1992: Visiting Scientist, Savannah River Ecology Lab, on sabbatical from VPI & SU.

1983: Visiting Scientist, Smithsonian Institution (National Zoological Park), on sabbatical from VPI & SU.

1971-1977: Assistant Professor, Department of Biology, VPI & SU.

1970-1971: Postdoctoral Research Associate, Harvard University.